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• All other flowering non-woody plants
• All other herbaceous, flowering dicots
• Podophyllum peltatum

Podophyllum peltatum — mandrake, May-apple

Facts

May-apple is distributed across the eastern United States, and reaches the northern limit of its distribution in New England, where it is rare. Its conservation is complicated by the fact that some populations are introduced. The ripe fruit is edible, but the rest of the plant is toxic. Some compounds that are found in the roots have been shown to have anti-tumor properties, and a derivitive of one of these compounds is used to treat some cancers.

Habitat

Anthropogenic (man-made or disturbed habitats), forest edges, forests, meadows and fields, shores of rivers or lakes

Characteristics

• Clonal plantlets

  Bulbils

  the plant does not appear to have bulbils

  Bulblets replace flowers

  there are no bulblets where the flowers are located

• Flowers

  Anther attachment

  the anther is attached by its base to the filament

  Anther color

  the anthers show no hint of a pink, reddish or purplish tint

  Anther length

  5–8 mm

  Anther opening

  the anthers have narrow slits or furrows that run lengthwise along the anthers

  Anther spurs

  the anthers do not have spurs on them

  Anther tube length

  0 mm

  Calyx growth after flowering

  the calyx does not grow to cover or partially cover the fruit

  Calyx symmetry

  there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)

  Carpel hairs

  the carpels have no hairs

  Carpels fused

  the carpel is solitary or (if 2 or more) the carpels are not fused to one another

  Cilia on petals

  the petal margins do not have cilia

  Cleistogamous flowers

  there are no cleistogamous flowers on the plan

  Corolla morphology

  NA

  Corolla palate

  no

  Corona lobe length

  0 mm

  Epicalyx

  the flower does not have an epicalyx

  Epicalyx number of parts

  0

  Filament length

  3–5 mm

  Filament surface

  the filament is smooth, with no hairs or scales

  Flower appearance

  the flowers appear after the leaves have appeared

  Flower description

  the flower has a superior ovary, and lacks a hypanthium

  Flower diameter

  30–50 mm

  Flower length

  30–55 mm

  Flower number

  1

  Flower orientation

  the flower bends downwards or hangs downwards

  Flower petal color

  • pink to red
  • white

  Flower reproductive parts

  the flower has both pollen- and seed-producing parts

  Flower symmetry

  there are two or more ways to evenly divide the flower (the flower is radially symmetrical)

  Flowers sunken into stem

  no

  Form of style

  • the style is knob-like at the tip, and unbranched
  • the style is lobed at the tip, and unbranched

  Fringed petal edges

  the petals are not fringed

  Fused stamen clusters

  NA

  Fusion of sepals and petals

  both the petals and sepals are separate and not fused

  Hairs on flower stalk

  the flower stalk has no hairs on it

  Hairs on inflorescence

  the axis of the inflorescence has no hairs on it

  Horns in hoods (Asclepias)

  NA

  Hypanthium

  the flower does not have a hypanthium

  Hypanthium length

  0 mm

  Inflorescence one-sided

  • NA
  • the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks

  Inner tepals (Rumex)

  NA

  Length of flower stalk

  15–60 mm

  Length of peduncle

  15–60 mm

  Marks on petals

  there are no noticeable marks on the petals

  Nectar spur

  the flower has no nectar spurs

  Number of branches in umbel

  0

  Number of carpels

  1

  Number of pistils

  1

  Number of sepals, petals or tepals

  • there are seven or more petals, sepals, or tepals in the flower
  • there are six petals, sepals, or tepals in the flower

  Number of styles

  2

  Ovary position

  the ovary is above the point of petal and/or sepal attachment

  Perianth shape

  the perianth is rotate (platter-shaped, the corolla flattened, circular, with nearly horizontally spreading lobes)

  Petal and sepal arrangement

  the flower includes two cycles of petal- or sepal-like structures

  Petal and sepal colors

  • pink to red
  • white

  Petal appearance

  the petals are thin and delicate, and pigmented (colored other than green or brown)

  Petal base

  the petal narrows gradually or does not narrow at the base

  Petal folding in bud

  the petals in bud are arranged in a cycle with edges overlapping like roof shingles (imbricate)

  Petal folds or pleats

  the petals of the flower do not have folds or plaits

  Petal glandular dots or scales

  no

  Petal hairs (Viola)

  NA

  Petal hairs on inner/upper surface

  there are no hairs on the inner/upper petal surface

  Petal length

  15–35 mm

  Petal length relative to sepals

  the petals are longer than the sepals

  Petal nectaries

  the petals have at least one nectary

  Petal number

  6–9

  Petal shape

  the petal outline is obovate (roughly egg-shaped, but with the widest point above the middle of the leaf blade)

  Petal tip shape

  the petal tip is rounded

  Petal tips (Cuscuta)

  NA

  Petal width

  10–25 mm

  Raceme attachment (Veronica)

  NA

  Reproductive system

  all the flowers have both carpels and stamens (synoecious)

  Scales inside corolla

  no

  Sepal and petal color

  the sepals are different from the petals

  Sepal appearance

  the sepals are green or brown, and leaf-like in texture

  Sepal appendages

  the sepals do not have appendages on them

  Sepal appendages (Oenothera)

  NA

  Sepal auricles

  the sepals have no auricles

  Sepal cilia

  the sepals do not have cilia

  Sepal color

  • green to brown
  • white

  Sepal features

  the sepals do not have any of the mentioned special features

  Sepal length

  10–18 mm

  Sepal number

  6

  Sepal relative length

  NA

  Sepal shape

  the sepal outline is roughly circular, as wide as long

  Sepal texture

  the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture

  Sepal tip shape

  the sepal tip is rounded

  Sepal uniformity

  all the sepals are about the same size

  Sepals fused only to sepals

  the sepals are separate from one another

  Spur length

  0 mm

  Spur number

  NA

  Stamen appendages

  stamen appendages are absent

  Stamen attachment

  the stamens are not attached to the petals or tepals

  Stamen length

  8–13 mm

  Stamen lengths differ

  the stamens are all approximately the same length

  Stamen morphology

  the stamens within each cycle are the same

  Stamen number

  • 12
  • 13 or more

  Stamen position relative to petals

  NA

  Stamen relative length

  anything

  Stamens fused

  the stamens are not attached to one another

  Staminodes

  there are no staminodes on the flower

  Stigma length

  3–6 mm

  Stigma position

  the stigmas are positioned at the tip of the style

  Style length

  1–2 mm

  Style petal-like

  the styles are not petal-like

  Style relative length

  NA

  Surface of ovary

  the ovary surface has no points, bumps or wrinkles

  Umbel flower reproductive parts

  NA

  Upper lip of bilabiate corolla

  NA

• Fruits or seeds

  Achene relative orientation

  NA

  Achene shape

  NA

  Achene surface (Polygonum)

  NA

  Achene type

  NA

  Berry color

  • orange
  • other
  • red
  • yellow

  Capsule color (Viola)

  NA

  Capsule ribs

  NA

  Capsule splitting

  NA

  Carpel beak length

  0 mm

  Fruit (pyxis) dehiscence

  NA

  Fruit beak length

  0 mm

  Fruit cross-section

  the fruit is round in cross-section

  Fruit features (Brassicaceae)

  NA

  Fruit length

  35–55 mm

  Fruit length relative to sepals

  NA

  Fruit length to width ratio

  1.4–1.8

  Fruit locules

  one

  Fruit shape

  • the fruit is ellipsoid (widest in the middle and tapering to each end)
  • the fruit is ovoid (egg-shaped)

  Fruit stalk orientation

  the fruits curve or droop downwards

  Fruit type (general)

  the fruit is fleshy

  Fruit type (specific)

  the fruit is a berry (fleshy, with the wall enclosing one or more sections, with two or more seeds)

  Fruit width

  20–40 mm

  Hair type on fruit

  NA

  Hairs on fruit

  the fruits are not hairy

  Legumes (Fabaceae)

  NA

  Mericarp length

  0 mm

  Mericarp segment shape (Desmodium)

  NA

  Other markings on berry

  the ripe berries are mostly one color without spots or streaks

  Ovary stipe

  the ovary or fruit does not have a stipe

  Placenta arrangement

  marginal

  Relative fruit length

  0

  Rows of seeds in fruit (Brassicaceae)

  NA

  Schizocarpic fruit compression

  NA

  Schizocarpic fruit segments

  0

  Seed length

  6–8 mm

  Seed number

  30–50

  Seed relative length

  the seed is longer than it is wide

  Seeds comose

  no hairs

  Septum in fruit (Brassicaceae)

  NA

  Wings on fruit

  the fruit does not have wings on it

  prickles on fruits

  the fruits do not have thorn-like defensive structures

• Glands or sap

  Glands on leaf blade

  the leaf blades do not have glandular dots or scales

  Sap

  the sap is clear and watery

  Sap color

  the sap is clear

• Growth form

  Growth form

  the plant is an herb (it has self-supporting stems)

  Horizontal rooting stem

  the plant does not have stolons

  Lifespan

  the plant lives more than two years

  Parasitism

  the plant is not parasitic

  Plant color

  the leaves or young stems of the plant are green

  Plants darken when dry

  no

  Spines on plant

  the plant has no spines

  Underground organs

  the plant has a rhizome (a horizontal underground stem with roots growing from it)

• Leaves

  Bracteole edges

  NA

  Bracteole length

  0 mm

  Bracteole number (Apiaceae)

  0

  Bracteole shape

  NA

  Bracteoles

  there are no bracteoles on the plant

  Bracts in plantain (Plantago)

  NA

  Final leaf segment length (compound lvs only)

  0 mm

  Final leaf segment length to width ratio (compound lvs only)

  0

  Final leaf segment width (compound lvs only)

  0 mm

  Floral bract texture

  the floral bracts are green, with a leaf-like texture

  Floral bracts

  • the flower does not have bracts
  • the flower has one or more bracts associated with it

  Hairs on leaf stalk

  • the petiole has hairs on it
  • the petiole has no hairs on it

  Hairs on underside of leaf

  the underside of the leaf is not hairy, or it has very few hairs

  Hairs on upper side of leaf

  the upper side of the leaf is not hairy, or it has very few hairs

  Hooked hairs on underside of leaf

  no

  Inflated hairs on leaf

  the leaf blade does not have inflated hairs on it

  Leaf arrangement

  • alternate: there is one leaf per node along the stem
  • opposite: there are two leaves per node along the stem

  Leaf blade base

  the leaf has a distinct leaf stalk (petiole)

  Leaf blade base shape

  the base of the leaf blade is rounded

  Leaf blade base symmetry

  the leaf blade base is symmetrical

  Leaf blade bloom

  the underside of the leaf has no noticeable bloom

  Leaf blade edges

  the edge of the leaf blade has lobes, or it has both teeth and lobes

  Leaf blade flatness

  • the edges of the leaf are curled under
  • the leaf is flat (planar) at the edges

  Leaf blade hairs

  the leaf blade has simple hairs with no glands, and not tangled or wooly

  Leaf blade length

  60–380 mm

  Leaf blade primary vein pattern

  the major veins radiate out from one point at the base

  Leaf blade shape

  • the leaf blade is orbicular (roughly circular, as wide as long)
  • the leaf blade is reniform (kidney-shaped; wider than long)

  Leaf blade surface colors

  the upper side of the leaf blade is relatively uniform in color

  Leaf blade texture

  the leaf blade is herbaceous (has a leafy texture)

  Leaf blade vein pattern

  the major veins of the leaf blade branch, but do not rejoin

  Leaf blade veins

  NA

  Leaf blade width

  100–400 mm

  Leaf duration

  the leaves drop off in winter (or they whither but persist on the plant)

  Leaf form

  the leaves are green, with an expanded blade and a leaf-like texture

  Leaf hair orientation

  the hairs are standing up straight or curved in different directions

  Leaf sheath length

  0 mm

  Leaf shiny

  the upper side of the leaf is dull or slightly shiny

  Leaf spines

  there are no spines on the leaf edges

  Leaf stalk

  the leaves have leaf stalks

  Leaf stalk attachment to leaf

  the petiole attaches at the center of the leaf blade (like an umbrella), or somewhere between the edge and the center

  Leaf stalk base

  the petiole base is narrow where it attaches to the stem

  Leaf stalk length

  50–150 mm

  Leaf teeth and lobes

  • the leaf blade margin has outward-pointing teeth
  • the leaf has lobes that radiate from the base, somewhat like a hand

  Leaf tip

  the tip of the leaf blade is rounded, with no point

  Leaf tufts in axils

  there are no clusters of smaller leaves growing out of axils

  Leaf type

  the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)

  Leaf types

  • NA
  • There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).

  Leaf variation

  • the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
  • the lower leaves are larger, toothier, and/or on longer stalks than the upper leaves

  Leaflet number

  0

  Leaflet petiolules

  NA

  Leaves per node

  • there are two leaves per node along the stem
  • there is one leaf per node along the stem

  Pinnately compound leaf type

  NA

  Specific leaf type

  the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)

  Stipels

  NA

  Stipule edges

  NA

  Stipule features

  NA

  Stipule fused to leaf stalk

  NA

  Stipule length

  0 mm

  Stipule shape

  NA

  Stipules

  there are no stipules on the plant

• Place

  Habitat

  • terrestrial
  • wetlands

  New England state

  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont

  Specific habitat

  • edges of forests
  • forests
  • man-made or disturbed habitats
  • meadows or fields
  • shores of rivers or lakes

• Stem, shoot, branch

  Branched tendrils

  NA

  Direction of stem hairs

  NA

  Flowering stem cross-section

  the flowering stem is circular, or with lots of small angles so that it is roughly circular

  Hair between stem nodes

  the stem has no hairs between the nodes

  Hairs between stem nodes

  the stem has no hairs between the nodes

  Hooked hairs on stem between nodes

  no

  Leaves on stem

  there is at least one full leaf above the base of the flowering stem

  Length of hairs between stem nodes

  0 mm

  Plant height

  20–60 cm

  Stem bloom

  there is no powdery or waxy film on the stem

  Stem hair distribution

  NA

  Stem nodes swollen

  the stem is not swollen at the nodes

  Stem orientation

  the stems are upright or angled outwards

  Stem roughness between nodes

  the stem does not feel rough

  Stem spacing

  the plant is solitary, or a few plants are growing together

  Stem succulence

  the stems are not succulent

  Tendril origin

  NA

  Tendrils

  the plant does not have tendrils

  Wings on stem

  the stem does not have wings on it

Wetland status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

New England distribution and conservation status

From the dichotomous key of Flora Novae Angliae