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Isoetes hieroglyphica — carved quillwort

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New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Facts

Carved quillwort is found mainly in the eastern half of Canada. It is very rare in New England, where it is known only from cold, clear water of lakes and slow-moving streams on gravel or cobble.

Habitat

Lacustrine (in lakes or ponds), riverine (in rivers or streams)

Characteristics

Habitat
aquatic
New England state
Maine
Leaf shape
the vegetative leaves are long and narrow
Spore leaf arrangement
the sporophylls are located in a whorl at the base of the plant
Form of shoot
NA
Horizontal stem
NA
Leaf differences
the vegetative leaves within a node are all similar in size and shape
Teeth on leaf edges
the edges of the vegetative leaves have no teeth
Spore leaf length
Up to 250 mm
Leaf outline
the vegetative leaves are long and very narrow (linear)
Show all characteristics
  • Clonal plantlets

    Gemma arrangement
    NA
    Gemma shape
    NA
    Gemma width
    0 mm
  • Leaves

    Leaf differences
    the vegetative leaves within a node are all similar in size and shape
    Leaf length
    Up to 250 mm
    Leaf orientation
    • the vegetative leaves spread away from the stem
    • the vegetative leaves spread slightly away from the stem, at a steep angle
    Leaf outline
    the vegetative leaves are long and very narrow (linear)
    Leaf ranks
    NA
    Leaf shape
    the vegetative leaves are long and narrow
    Pores on leaves
    there are no apparent pores on the vegetative leaves
    Spore leaf length
    Up to 250 mm
    Teeth on leaf edges
    the edges of the vegetative leaves have no teeth
  • Place

    Habitat
    aquatic
    New England state
    Maine
    Specific habitat
    • in lakes or ponds
    • in rivers or streams
  • Spores or spore cones

    Cone base at stem
    NA
    Cone stalk branching
    NA
    Cone thickness
    0 mm
    Cone width
    0 mm
    Length of cone
    0 mm
    Number of cones
    0
    Quillwort itssue covering spores
    the velum covers less than half of the spores
    Same or different spores
    there are two different types of spores present
    Spore diameter
    0.55–0.7
    Spore girdle
    there is a smooth girdle alongside the equatorial ridge
    Spore leaf arrangement
    the sporophylls are located in a whorl at the base of the plant
    Spore leaf lifespan
    the sporophylls remain green for the life of the plant
    Spore leaf orientation
    the sporophylls slant upwards at a steep angle
    Spore leaf shape
    the spore-bearing leaves are long and narrow
    Spore leaf teeth
    The edges of the spore-bearing leaves are smooth, and without teeth
    Spore texture
    the spore surface has a net-like pattern on it (reticulate)
    Sporophyll ranks
    NA
    Sterile tip of cone
    NA
  • Stem, shoot, branch

    Branch cross-section
    NA
    Branch form
    NA
    Constriction zones
    • NA
    • NA
    Form of shoot
    NA
    Horizontal stem
    NA
    Horizontal stem length
    0 mm
    Horizontal stem thickness
    0 mm
    Stem height
    0 mm

Wetland status

Occurs only in wetlands. (Wetland indicator code: OBL)

New England distribution and conservation status

Distribution

Connecticut
absent
Maine
present
Massachusetts
absent
New Hampshire
absent
Rhode Island
absent
Vermont
absent

Conservation status

None

Native to North America?

Yes

Sometimes confused with

Isoetes acadiensis:
megaspores megaspores 0.4–0.65 mm in diameter, averaging less than 0.6 mm, and leaves gradually tapering to the tip (vs. I. hieroglyphica, with megaspores 0.55–0.7 mm in diameter, averaging more than 0.6 mm, and leaves abruptly tapering to the tip).
Isoetes lacustris:
megaspores with a papillate girdle, cristate to reticulate with sharp or roughened crests (vs. I. hieroglyphica, with megaspores with a smooth girdle, reticulate with low, rounded ridges).

Synonyms

  • Isoetes lacustris L. forma hieroglyphica (A.A. Eat.) N.E. Pfeiff.

Family

Isoetaceae

Genus

Isoetes

From the dichotomous key of Flora Novae Angliae

4.  Isoetes hieroglyphica A.A. Eat. NC

carved quillwort. Isoetes lacustris L. forma hieroglyphica (A.A. Eat.) N.E. Pfeiff. • ME; most collections from the northern portion of ME. Cold, clear water of lakes and slow-moving streams with gravel and cobble substrate.