Linaria vulgaris P. Mill.

butter-and-eggs toadflax

Copyright: various copyright holders. To reuse an image, please click it to see who you will need to contact.

New England Distribution

Adapted from BONAP data

Found this plant? Take a photo and post a sighting.

North America Distribution

Adapted from BONAP data

enlarge

Facts About

The unusual name of butter-and-eggs comes from the yellow and cream colors of the flower of this invasive perennial. Each plant can produce up to thirty thousand winged seeds per year, and once established it is difficult to remove. Biological control may be helpful, using two weevil species (Gymnetron antirrhini and Mecinus janthinu), and two moths (Calophasia lunula and Eteobalea serratella).

Habitat

Anthropogenic (man-made or disturbed habitats), meadows and fields

Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
  • orange
  • yellow
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
  • alternate: there is one leaf per node along the stem
  • whorled: there are three or more leaves per node along the stem
Leaf blade edges
the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
5–11 mm
Show All Characteristics
  • Clonal plantlets
    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located
  • Flowers
    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla morphology
    the flower has two prominent lips
    Corolla palate
    yes
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower length
    19–33 mm
    Flower number
    5–30
    Flower petal color
    • orange
    • yellow
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is knob-like at the tip, and unbranched
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Hairs on flower stalk
    the flower stalk has hairs on it
    Hairs on inflorescence
    • at least some of the hairs on the axis of the inflorescence have glands
    • the axis of the inflorescence has no hairs on it
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Hypanthium length
    0 mm
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Length of flower stalk
    2–8 mm
    Marks on petals
    there are no noticeable marks on the petals
    Nectar spur
    the flower has one or more nectar spurs
    Number of branches in umbel
    0
    Number of carpels
    2
    Number of pistils
    1
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    the perianth is cylindric (tube-shaped)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • orange
    • yellow
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal hairs on inner/upper surface
    there are hairs on the inner/upper petal surface
    Petal length
    19–33 mm
    Petal length relative to sepals
    the petals are longer than the sepals
    Petal number
    5
    Petal shape
    the petal outline is ovate (widest below the middle and broadly tapering at both ends)
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal color
    green to brown
    Sepal length
    3–6 mm
    Sepal number
    5
    Sepal shape
    • the sepal outline is oblanceolate (lance-shaped, but with the widest point above the middle of the leaf blade)
    • the sepal outline is ovate (widest below the middle and broadly tapering at both ends)
    Sepal tip shape
    the sepal tip is acute (is sharply pointed)
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Spur length
    10–13 mm
    Spur number
    1
    Stamen lengths differ
    the stamens are didynamous (two long stamens and two short ones)
    Stamen morphology
    the stamens within a cycle differ in length or width
    Stamen number
    4
    Stamen relative length
    anything
    Style length
    7–9 mm
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    the upper lip of the bilabiate corolla has two lobes
  • Fruits or seeds
    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    the capsule has no prominent ribs or wings
    Capsule splitting
    the capsule splits by two main valves, teeth or pores
    Carpel beak length
    0 mm
    Fruit (pyxis) dehiscence
    NA
    Fruit features (Brassicaceae)
    NA
    Fruit length
    5–11 mm
    Fruit locules
    two
    Fruit shape
    • the fruit is obloid (longer than wide and with rounded ends)
    • the fruit is ovoid (egg-shaped)
    • the fruit is spherical
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
    Fruit width
    5–7 mm
    Hair type on fruit
    NA
    Hairs on fruit
    the fruits are not hairy
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Seed length
    1.5–4 mm
    Seed relative length
    the seed is about as long as it is wide
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures
  • Glands or sap
    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear
  • Growth form
    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    the plant has a rhizome (a horizontal underground stem with roots growing from it)
  • Leaves
    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    the flower has one or more bracts associated with it
    Hairs on leaf stalk
    NA
    Hairs on underside of leaf
    the underside of the leaf is not hairy, or it has very few hairs
    Hairs on upper side of leaf
    the upper side of the leaf is not hairy, or it has very few hairs
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    • alternate: there is one leaf per node along the stem
    • whorled: there are three or more leaves per node along the stem
    Leaf blade base
    the leaf has no stalk
    Leaf blade base shape
    the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade flatness
    the leaf is flat (planar) at the edges
    Leaf blade hairs
    NA
    Leaf blade length
    20–60 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is elliptic (widest near the middle and tapering at both ends)
    • the leaf blade is linear (very narrow with more or less parallel sides)
    • the leaf blade is oblanceolate (lance-shaped, but with the widest point above the middle of the leaf blade)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is herbaceous (has a leafy texture)
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    1–15 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf hair orientation
    NA
    Leaf sheath length
    0 mm
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have no leaf stalks, but attach directly to the stem
    Leaf stalk attachment to leaf
    NA
    Leaf stalk base
    NA
    Leaf stalk length
    0 mm
    Leaf teeth and lobes
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf tip
    the tip of the leaf blade is acute (sharply pointed)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    • there are four leaves per node along the stem
    • there is one leaf per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant
    Teeth per side of leaf blade
    0
  • Place
    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • man-made or disturbed habitats
    • meadows or fields
  • Scent
    Plant odor
    the plant does not have much of a smell
  • Stem, shoot, branch
    Branched tendrils
    NA
    Flowering stem cross-section
    the flowering stem is circular, or with lots of small angles so that it is roughly circular
    Hair between stem nodes
    • the stem has hairs between the nodes
    • the stem has no hairs between the nodes
    Hairs between stem nodes
    • at least some of the hairs on the stem have glands
    • the stem has no hairs between the nodes
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    15–90 cm
    Stem bloom
    • the stem has a powdery or waxy film on it that can be rubbed away
    • there is no powdery or waxy film on the stem
    Stem hair distribution
    the hairs on the stem are distributed more of less uniformly
    Stem nodes swollen
    the stem is not swollen at the nodes
    Stem orientation
    the stems are upright or angled outwards
    Stem roughness between nodes
    the stem does not feel rough
    Stem succulence
    the stems are not succulent
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland Status

Not classified

New England Distribution and Conservation Status

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation Status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
unranked (S-rank: SNR)

Native to North America?

No

Sometimes Confused With

Linaria spartea:
stigma bifid, seeds ovoid-pyramidal, unwinged, 0.6–0.7 mm long, and leaf blades 0.5–1 mm wide (vs. L. vulgaris, with the stigma capitate, seeds discoid, with a narrow, marginal wing, mostly 1.5–3 mm long, and leaf blades mostly 1–5 mm wide).

Synonyms

  • Antirrhinum orontium L.
  • Linaria linaria (L.) Karst.

Genus

Linaria

Need Help?

Get Help

Information from Dichotomous Key of Flora Novae Angliae

7.  Linaria vulgaris P. Mill. E

butter-and-eggs toadflax. Linaria linaria (L.) Karst. • CT, MA, ME, NH, RI, VT. Fields, roadsides, waste areas, railroads.