Polygala ambigua Nutt.

alternate milkwort

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New England Distribution

Adapted from BONAP data

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North America Distribution

Adapted from BONAP data

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Facts About

Alternate milkwort is widespread in New England but rare, occupying anthropogenically disturbed sites over much of New England, while growing in relatively pristine habitats in Connecticut. Its scientific name, "ambigua" may refer to its "ambiguous" status as a species; it was once thought to be a variety of the whorled milkwort (P. verticillata), but is now distinguished from that species by its green-white to white flowers and the fact that the upper nodes usually only have one leaf as opposed to a whorl of leaves.

Habitat

Anthropogenic (man-made or disturbed habitats), forest edges, meadows and fields, woodlands

Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
  • green to brown
  • white
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
alternate: there is one leaf per node along the stem
Leaf blade edges
the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
  • there are five petals, sepals, or tepals in the flower
  • there are three petals, sepals, or tepals in the flower
Fusion of sepals and petals
  • both the petals and sepals are separate and not fused
  • the petals or the sepals are fused into a cup or tube
Stamen number
8
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
1.3–1.7 mm
Show All Characteristics
  • Clonal plantlets
    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located
  • Flowers
    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there is only one way to evenly divide the calyx (the calyx is bilaterally symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower petal color
    • green to brown
    • white
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is lobed at the tip, and unbranched
    Fused stamen clusters
    there is one cluster of fused stamens
    Fusion of sepals and petals
    • both the petals and sepals are separate and not fused
    • the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inflorescence width
    2–14 mm
    Inner tepals (Rumex)
    NA
    Nectar spur
    the flower has no nectar spurs
    Number of pistils
    1
    Number of sepals, petals or tepals
    • there are five petals, sepals, or tepals in the flower
    • there are three petals, sepals, or tepals in the flower
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • green to brown
    • white
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal number
    3
    Petal tips (Cuscuta)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal number
    5
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen number
    8
    Stamen position relative to petals
    NA
    Stamens fused
    the stamens are attached to one another at or near their bases
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA
  • Fruits or seeds
    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    the capsule has no prominent ribs or wings
    Capsule splitting
    the capsule splits by two main valves, teeth or pores
    Fruit (pyxis) dehiscence
    NA
    Fruit beak length
    0 mm
    Fruit features (Brassicaceae)
    NA
    Fruit length
    1.3–1.7 mm
    Fruit locules
    two
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    • the plant has apical placentation, where one or a few ovules develop at the top of a simple or compound ovary
    • the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures
  • Glands or sap
    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear
  • Growth form
    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives only a single year or less
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    there is a thickened taproot on the plant
  • Leaves
    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    • the flower does not have bracts
    • the flower has one or more bracts associated with it
    Hairs on underside of leaf
    the underside of the leaf is not hairy, or it has very few hairs
    Hairs on upper side of leaf
    the upper side of the leaf is not hairy, or it has very few hairs
    Leaf arrangement
    alternate: there is one leaf per node along the stem
    Leaf blade base
    the leaf has no stalk
    Leaf blade base shape
    the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade edges
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade length
    10–20 mm
    Leaf blade shape
    the leaf blade is linear (very narrow with more or less parallel sides)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade width
    1–5 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have no leaf stalks, but attach directly to the stem
    Leaf teeth and lobes
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaves per node
    • there are five leaves per node along the stem
    • there are four leaves per node along the stem
    • there are three leaves per node along the stem
    • there are two leaves per node along the stem
    • there is one leaf per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule features
    NA
    Stipules
    there are no stipules on the plant
  • Place
    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • edges of forests
    • man-made or disturbed habitats
    • meadows or fields
    • woodlands
  • Scent
    Plant odor
    the plant does not have much of a smell
  • Stem, shoot, branch
    Branched tendrils
    NA
    Flowering stem cross-section
    • the flowering stem has strong or sharp angles, but is not square
    • the flowering stem is roughly square
    Hair between stem nodes
    the stem has no hairs between the nodes
    Hairs between stem nodes
    the stem has no hairs between the nodes
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    10–40 cm
    Stem orientation
    the stems are upright or angled outwards
    Stem spacing
    the plant is solitary, or a few plants are growing together
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils

Wetland Status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

New England Distribution and Conservation Status

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation Status

Exact status definitions can vary from state to state. For details, please check with your state.

Connecticut
unrankable (S-rank: SU), special concern (code: SC)
New Hampshire
unrankable (S-rank: SU), Ind (code: Ind)
Rhode Island
historical (S-rank: SH), state historical (code: SH)
Vermont
extremely rare (S-rank: S1)

Native to North America?

Yes

Sometimes Confused With

Polygala verticillata

Synonyms

  • Polygala verticillata L. var. ambigua (Nutt.) Wood
  • Polygala verticillata L. var. dolichoptera Fern.

Family

Polygalaceae

Genus

Polygala

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Information from Dichotomous Key of Flora Novae Angliae

1.  Polygala ambigua Nutt. NC

alternate milkwort. Polygala verticillata L. var. ambigua (Nutt.) Wood; P. verticillata L. var. dolichoptera Fern. • CT, MA, ME, NH, RI, VT. Fields, forest edges, roadsides, railroads, woodlands, forest clearings. This species utilizes anthropogenic habitats over most of 
New England but has been collected from relatively pristine sites in CT.